November 17, 2012 at 7:03 PM

Indiana tested Penn State's secondary Saturday.

The Hoosiers threw the ball 59 times against the Nittany Lions, who were without starting safety Malcolm Willis. Penn State allowed quarterback Cameron Coffman to accumulate 454 passing yards, but the team's secondary bent and didn't break in a 22-point victory.

"I thought those guys were resilient, they are tough," coach Bill O'Brien said. "Maybe there is a bad play here and there, but they never let it become a bad day, if that makes sense."

Jake Fagnano started in place of Willis and safety Ryan Keiser got some playing time on defense for the first time in his career. Kesier made three tackles, as well did Fagnano who also broke up two passes.

Senior cornerback Stephon Morris said he thought the defensive backs held their own against a fast-paced Indiana attack.

"Fagnano got his first start today, did really well," Morris said. "Keiser, played some and he did good. That was his first time actually playing defense, so he stepped up."

Coffman was able to complete 33 of his attempts, but the secondary did force one turnover in the form of an Adrian Amos interception. It was the cornerback's second pick of the season, and he remains the only Nittany Lion defensive back with an interception in 11 games.