MADISON, Wis. — Just like after his previous 343 collegiate tackles, Paul Posluszny was all business.

The Penn State linebacker eclipsed Greg Buttle’s career tackle record in Saturday’s loss at Wisconsin, but the humble senior’s joy over the accomplishment wasn’t just hidden — it was essentially nonexistent.

"In this situation where it’s a huge game and we lost, it means something," Posluszny said of securing the record. "But I would much rather have the win."

Posluszny passed Buttle’s mark on a third-and-5 play in the second quarter. Wisconsin quarterback John Stocco dumped a pass to tight end Travis Beckum, who gained four yards before being stopped short on the record-breaking tackle, Posluszny’s ninth of the opening half.

Given the magnitude of the game and the co-captain’s typical intensity, Posluszny had no idea he’d become the all-time statistical leader at "Linebacker U" until someone told him 30 minutes after the game.

Defensive coordinator Tom Bradley, who played with Buttle at Penn State, was also unaware of the feat until it was brought to his attention following the game.

"It’s quite an accomplishment," Bradley said. "Paul’s the heart and soul of our defense, and it couldn’t have happened to a more deserving, finer young man than Paul Posluszny. That’s a record that will take a long time to break. He’s a super person, super football player, and I’m proud to have him on my defense."

Posluszny finished the game with 14 takedowns, the most by any Nittany Lion this season and the most he’s had since last year’s Oct. 8 game against Ohio State.

At the start of this year, some had questioned Posluszny’s health after off-season knee rehabilitation and a position change from outside linebacker to middle.

While the reigning Butkus and Bednarik Award winner, as well as fellow linebackers Dan Connor and Sean Lee, played well, the defense still had its hands full with Wisconsin’s sizeable offensive line and tailback P.J. Hill, who rushed for 148 yards on 31 carries.

"They got a lot of yards on us," Posluszny said. "We thought we would play the run better than we did. They came out and ran the ball well, and we’re a little disappointed with that defensively."

Though obviously frustrated by the game’s outcome and the slew of injuries to his teammates and Joe Paterno, Posluszny — fighting through his usual modesty — admitted that someday the record will be more significant to him.

"It’ll probably mean something later on, once I’m done," Posluszny said. "I wish I had it under better circumstances. The win was the most important thing here and we didn’t get it today."